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J Herbmed Pharmacol. 2026;15(1): 75-81.
doi: 10.34172/jhp.2026.53309
  Abstract View: 7
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Original Article

Antiproliferative and apoptosis-inducing effects of Peganum harmala L. seed extracts on gastric adenocarcinoma

Mohammad-Taghi Moradi 1 ORCID logo, Nafiseh Bagherian Khouzani 2 ORCID logo, Ali Rafieian 3 ORCID logo, Majid Asadi-Samani 4* ORCID logo, Mohammad Reza Khosravi Farsani 5 ORCID logo, Niloufar Mousiany 6 ORCID logo

1 Medical Plants Research Center, Basic Health Sciences Institute, Shahrekord University of Medical Sciences, Shahrekord, Iran
2 Department of Pharmaceutical Chemistry, College of Pharmacy, Al-Zahraa University for Women, Karbala, Iraq
3 Student Research Center, Faculty of Medicine, Shahid Beheshti University of Medical Sciences, Tehran, Iran
4 Cellular and Molecular Research Center, Basic Health Sciences Institute, Shahrekord University of Medical Sciences, Shahrekord, Iran
5 Clinical Chemistry Research Center, Basic Health Sciences Institute, Shahrekord University of Medical Sciences, Shahrekord, Iran
6 Student Research Committee, Shahrekord University of Medical Sciences, Shahrekord, Iran
*Corresponding Author: Majid Asadi-Samani, Email: asadi.m@skums.ac.ir

Abstract

Introduction: Gastric cancer treatment remains challenging due to side effects and limited options. This study investigated Peganum harmala L., a traditional medicinal plant with anticancer alkaloids, by evaluating its crude extract and fractions for antiproliferative, apoptotic, and cell cycle effects on human gastric adenocarcinoma (AGS) cells.

Methods: Fractionation of the crude hydroalcoholic extract was done using solvents of varying polarity (chloroform, ethyl acetate, n-butanol, and n-hexane). Antiproliferative effects were assessed via MTT assay at five concentrations (0-200 µg/mL) on AGS and human dermal fibroblast (HDF) cells. Apoptosis and cell cycle were conducted via flow cytometric technique after 48 h treatment by annexin V-FITC/propidium iodide binding and propidium iodide intercalation into nuclear DNA, respectively.

Results: The crude extract and three fractions (chloroform, n-butanol, ethyl acetate) showed selective cytotoxicity against AGS cells versus HDFs (P<0.05). The n-butanol fraction indicated the highest potency compared to the crude extract and other fractions (P<0.05). Apoptosis analysis via flow cytometry indicated a noticeable increase in apoptosis in AGS cells treated with P. harmala crude extract, compared to untreated cells (P<0.05). Flow cytometric analysis revealed that both the crude extract and n-butanol fraction significantly arrested cells in the G2/M phase while reducing the G1 phase population (P<0.05).

Conclusion: P. harmala seed extracts demonstrate significant, selective antiproliferative effects on gastric cancer cells, mediated partly through the induction of apoptosis and arresting the cell cycle in G2/M phase. These findings validate traditional uses and highlight the n-butanol fraction as particularly promising for further anticancer development.


Implication for health policy/practice/research/medical education:

Peganum harmala seed extract selectively targets gastric cancer cells with minimal harm to normal cells. Among the crude extract and other fractions, the n-butanol fraction showed the strongest activity, inducing apoptosis and cell cycle arrest. This suggests it is a promising natural adjuvant therapy with fewer side effects than conventional treatments.

Please cite this paper as: Moradi MT, Bagherian Khouzani N, Rafieian A, Asadi-Samani M, Khosravi Farsani MR, Mousiany N. Antiproliferative and apoptosis-inducing effects of Peganum harmala L. seed extracts on gastric adenocarcinoma. J Herbmed Pharmacol. 2026;15(1):75-81. doi: 10.34172/jhp.2026.53309.

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Submitted: 05 May 2025
Revision: 05 Aug 2025
Accepted: 25 Aug 2025
ePublished: 01 Jan 2026
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